Dr. Julia Ann Schultz

Functional morphology of the early mammalian dentition and jaw biomechanics, reconstruction of mastication pattern, striation analysis, facet formation and wear pattern / Evolution of the mammalian middle and inner ear / Mammalian diversity and disparity in the Mesozoic / Micro-computed tomography, 3D-reconstruction and virtual simulation

Consultation hours

By appointment

Research topics

  • Mammal evolution
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Dr. Julia Ann Schultz

Pal 1.001

Nussallee 8

53115 Bonn

Vertebrate - Mammalian

Curiculum vitae

  • Current position: Research Assistant at the Institute of Geosciences, Universität Bonn
    2015-2017 DAAD postdoc fellow in the Luo Lab at The University of Chicago, Department of Organismal Biology and Anatomy. https://luolab.uchicago.edu/people/

  • 2011-2015 Postdoc, Institute of Geosciences, Universität Bonn: project D1 within DFG research unit 771 “Function and performance enhancement in the mammalian feeding apparatus” (second funding period). Link: https://tinyurl.com/4m85yszm

  • Nov. 2011 Doctoral-Thesis defense: Funktionelle Morphologie und Abnutzungsmuster prätribosphenischer Molaren am Beispiel der Dryolestida (Mammalia, Cladotheria). Link: https://tinyurl.com/mr34wu4w

  • 2008-2011 PhD student, Institute of Geosciences (former Steinmann-Institut für Geologie, Mineralogie und Paläontologie), Universität Bonn: project D1 of the DFG research unit 771 “Function and performance enhancement in the mammalian dentition” (first funding period). Link: https://tinyurl.com/4m85yszm

  • 1998-2007 Study of Geology, Freie Universität Berlin, degree: Dipl.-Geol., Diploma-Thesis: The Inner Ear Morphology throughout the Mammalian Evolution - A Comparative Study

Selected publications

Martin,T. and Schultz,J.A. 2023. „Deciduous dentition, tooth replacement, and mandibular growth in the Late Jurassic docodontan Haldanodon exspectatus (Mammaliaformes)“. Journal of Mammalian Evolution.
https://doi.org/10.1007/s10914-023-09668-2
Keppeler,H., Schultz,J.A., Ruf,I. and Martin,T. 2023. „Cranial anatomy of Hypisodus minimus (Artiodactyla: Ruminantia) from the Oligocene Brule Formation of North America“. Palaeontographica Abteilung A. 327(1-3): 55-92.
https://doi.org/10.1127/pala/2023/0140
Schwermann,A.H., Schultz,J.A., Fuchs,E., Leha,A., Hummel,J., Kullmer,O., Steuer,P. and Martin,T. 2023. „Does age matter?—Efficiency of mechanical food break down in Tupaia belangeri at different ages“. PLOS ONE. 18(7): e0274439.
https://doi.org/10.1371/journal.pone.0274439
Martin,T., Averianov,A.O., Schultz,J.A. and Schwermann,A.H. 2023. „A stem therian mammal from the Lower Cretaceous of Germany“. Journal of Vertebrate Paleontology.
https://doi.org/10.1080/02724634.2023.2224848
Martin,T., Goin,F.J., Schultz,J.A. and Gelfo,J.N. 2022. „Early Late Cretaceous mammals from southern Patagonia (Santa Cruz province, Argentina)“. Cretaceous Research. 133: 105127.
https://doi.org/10.1016/j.cretres.2021.105127
Martin,T., Averianov,A., Schultz,J., Schellhorn,R. and Schwermann,A. 2022. „First spalacotheriid and dryolestid mammals from the Cretaceous of Germany“. Acta Palaeontologica Polonica. 67:
https://doi.org/10.4202/app.00914.2021
Schultz,J.A., Schellhorn,R., Skutschas,P.P., Vitenko,D.D., Kolchanov,V.V., Grigoriev,D.V., Kuzmin,I.T., Kolosov,P.N., Lopatin,A.V., Averianov,A.O. and Martin,T. 2022. „Mammalian petrosal from the Lower Cretaceous high paleo-latitude Teete locality (Yakutia, Eastern Russia)“. Vertebrate Zoology. 72: 159-168.
https://doi.org/10.3897/vz.72.e78479
Begat,A., Kriwet,J., Gelfo,J.N., Cavalli,S.G., Schultz,J.A. and Martin,T. 2022. „The first southern hemisphere occurrence of the extinct Cretaceous sclerorhynchoid sawfish Ptychotrygon (Chondrichthyes, Batoidea), with a review of Ptychotrygon taxonomy“. Journal of Vertebrate Paleontology. 42(2):
https://doi.org/10.1080/02724634.2022.2162411
Schultz,J.A., Ruf,I., Averianov,A.O., Schellhorn,R., Lopatin,A.V. and Martin,T. 2022. „Jurassic mammaliaform petrosals from Western Siberia (Russia) and implications for early mammalian inner-ear anatomy“. Zoological Journal of the Linnean Society. 196(3): 1175-1200.
https://doi.org/10.1093/zoolinnean/zlab096
Grossnickle,D.M., Weaver,L.N., Jäger,K.R.K. and Schultz,J.A. 2022. „The evolution of anteriorly directed molar occlusion in mammals“. Zoological Journal of the Linnean Society. 194(2): 349-365.
https://doi.org/10.1093/zoolinnean/zlab039
Martin, T., O. Averianov, A., A. Schultz, J., H. Schwermann, A. and Wings, O. 2021. „Late Jurassic multituberculate mammals from Langenberg Quarry (Lower Saxony, Germany) and palaeobiogeography of European Jurassic multituberculates“. Historical Biology. 33(5): 616-629.
https://doi.org/10.1080/08912963.2019.1650274
Schultz, J. A., U. Anders, C. Braune, J. Brinkkötter, I. Calandra, S. Engels, E. Findeisen, J.-P. Gailer, J. Hummel, K. R. K. Jäger, T. M. Kaiser, D. C. Kalthoff, W. v. Koenigswald, O. Kullmer, C. Landwehr, M. Mau, U. Menz, I. Ruf, A. M. Schubert, E. Schulz-Kornas, A. H. Schwermann, L. C. Schwermann, M. Skiba, P. Steuer, K.-H. Südekum, D. E. Winkler, T. Martin 2021. „A new facet terminology for mammalian dentitions“. In: Mammalian Teeth - Form and Function. pp 11-24. T. Martin & W. v. Koenigswald, eds.. München (Pfeil-Verlag). ISBN: 978-3-89937-266-3.
Schultz, J. A., S. Engels, L. C. Schwermann, W. v. Koenigswald 2021. „Evolutionary trends in the mastication patterns in some perissodactyls, cetartiodactyls, and proboscideans“. In: Mammalian Teeth - Form and Function. pp215-230. T. Martin & W. v. Koenigswald, eds.. München (Pfeil-Verlag). ISBN: 978-3-89937-266-3.
Martin, T., J J. Brinkkötter, K. R. K. Jäger, T. Plogschties, A. H. Schwermann, J. A. Schultz 2021. „Molar diversity and functional adaptations in Mesozoic mammals“. In: Mammalian Teeth - Form and Function. pp187-214. T. Martin & W. v. Koenigswald, eds.. München (Pfeil-Verlag). ISBN: 978-3-89937-266-3.
Martin,T., Averianov,A.O., Schultz,J.A., Schwermann,A.H. and Wings,O. 2021. „A derived dryolestid mammal indicates possible insular endemism in the Late Jurassic of Germany“. The Science of Nature. 108(3): .
https://doi.org/10.1007/s00114-021-01719-z
Martin,T., Averianov,A.O., Schultz,J.A. and Schwermann,A.H. 2021. „First multituberculate mammals from the Lower Cretaceous of Germany“. Cretaceous Research. 119(): 104699.
https://doi.org/10.1016/j.cretres.2020.104699
Averianov,A.O., Martin,T., Lopatin,A.V., Schultz,J.A., Schellhorn,R., Krasnolutskii,S., Skutschas,P. and Ivantsov,S. 2021. „Multituberculate mammals from the Middle Jurassic of Western Siberia, Russia, and the origin of Multituberculata“. Papers in Palaeontology. 7(2): 769-787.
https://doi.org/10.1002/spp2.1317
Skutschas, P., Kolchanov, V., Krasnolutskii, S., Averianov, A., Schellhorn, R., Schultz, J. and Martin, T. 2020. „A new small-sized stem salamander from the Middle Jurassic of Western Siberia, Russia“. PLOS ONE. 15(2): e0228610.
https://doi.org/10.1371/journal.pone.0228610
Krause,D.W., Hoffmann,S., Hu,Y., Wible,J.R., Rougier,G.W., Kirk,E.C., Groenke,J.R., Rogers,R.R., Rossie,J.B., Schultz,J.A., Evans,A.R., von Koenigswald,W. and Rahantarisoa,L.J. 2020. „Skeleton of a Cretaceous mammal from Madagascar reflects long-term insularity“. Nature. 581(7809): 421-427.
https://doi.org/10.1038/s41586-020-2234-8
Schultz,J.A. 2020. „Eat and listen—how chewing and hearing evolved“. Science. 367(6475): 244-246.
https://doi.org/10.1126/science.aba3808
Krause,D.W., Hu,Y., Hoffmann,S., Groenke,J.R., Schultz,J.A., Evans,A.R., von Koenigswald,W. and Rougier,G.W. 2020. „Dental morphology of Adalatherium hui (Mammalia, Gondwanatheria) from the LATE Cretaceous of Madagascar“. Journal of Vertebrate Paleontology. 40(sup1): 97-132.
https://doi.org/10.1080/02724634.2020.1811292
Averianov, A., Krasnolutskii, S., Ivantsov, S., Skutschas, P., Schellhorn, R., Schultz, J. and Martin, T. 2019. „Sauropod remains from the Middle Jurassic Itat Formation of West Siberia, Russia“. PalZ. 93(4): 691-701.
https://doi.org/10.1007/s12542-018-00445-8
Averianov, A.O., Martin, T., Lopatin, A.V., Schultz, J.A., Schellhorn, R., Krasnolutskii, S., Skutschas, P. and Ivantsov, S. 2019. „Haramiyidan mammals from the Middle Jurassic of Western Siberia, Russia. Part 1: Shenshouidae and Maiopatagium“. Journal of Vertebrate Paleontology. 39(4): e1669159.
https://doi.org/10.1080/02724634.2019.1669159
Engels, S. and Schultz, J.A. 2019. „Evolution of the power stroke in early Equoidea (Perissodactyla, Mammalia)“. Palaeobiodiversity and Palaeoenvironments. 99(2): 271-291.
https://doi.org/10.1007/s12549-018-0341-4
Kalthoff, D.C., Schulz-Kornas, E., Corfe, I., Martin, T., McLoughlin, S. and Schultz, J.A. 2019. „Complementary approaches to tooth wear analysis in Tritylodontidae (Synapsida, Mammaliamorpha) reveal a generalist diet“. PLOS ONE. 14(7): e0220188.
https://doi.org/10.1371/journal.pone.0220188
Panciroli, E., Schultz, J.A. and Luo, Z. 2019. „Morphology of the petrosal and stapes ofBorealestes(Mammaliaformes, Docodonta) from the Middle Jurassic of Skye, Scotland“. Papers in Palaeontology. 5(1): 139-156.
https://doi.org/10.1002/spp2.1233
Panciroli, E., Schultz, J.A. and Luo, Z.-X. 2019. „Data from: Morphology of the petrosal and stapes of Borealestes (Mammaliaformes, Docodonta) from the Middle Jurassic of Skye, Scotland“. Dryad.
https://doi.org/10.5061/dryad.07934k0
Pfaff, C., Schultz, J.A. and Schellhorn, R. 2019. „The vertebrate middle and inner ear: A short overview“. Journal of Morphology. 280(8): 1098-1105.
https://doi.org/10.1002/jmor.20880
Huttenlocker, A.K., Grossnickle, D.M., Kirkland, J.I., Schultz, J.A. and Luo, Z. 2018. „Late-surviving stem mammal links the lowermost Cretaceous of North America and Gondwana“. Nature. 558(7708): 108-112.
https://doi.org/10.1038/s41586-018-0126-y
Schultz, J.A., Ruf, I. and Martin, T. 2018. „Oldest known multituberculate stapes suggests an asymmetric bicrural pattern as ancestral for Multituberculata“. Proceedings of the Royal Society B: Biological Sciences. 285(1873): 20172779.
https://doi.org/10.1098/rspb.2017.2779
Averianov, A.O., Martin, T., Lopatin, A.V., Schultz, J.A., Skutschas, P.P., Rico, S. and Krasnolutskii, S.A. 2017. „A tritylodontid synapsid from the Middle Jurassic of Siberia and the taxonomy of derived tritylodontids“. Journal of Vertebrate Paleontology. 37(5): e1363767.
https://doi.org/10.1080/02724634.2017.1363767
Schultz, J. A., Bhullar B. A. S. & Luo, Z. X. 2017. „Re-examination of the Jurassic mammaliaform Docodon victor by computed tomography and occlusal functional analysis“. Journal of Mammalian Evolution. 1-30.
Schultz, J.A., Menz, U., Winkler, D.E., Schulz-Kornas, E., Engels, S., Kalthoff, D.C., Koenigswald, W.v. , Ruf, I., Kaiser, T.M., Kullmer, O., Südekum, K. and Martin, T. 2017. „Modular Wear Facet Nomenclature for mammalian post-canine dentitions“. Historical Biology. 30(1-2): 30-41.
https://doi.org/10.1080/08912963.2017.1302442
Ferner,K., Schultz,J.A. and Zeller,U. 2017. „Comparative anatomy of neonates of the three major mammalian groups (monotremes, marsupials, placentals) and implications for the ancestral mammalian neonate morphotype“. Journal of Anatomy. 231(6): 798-822.
https://doi.org/10.1111/joa.12689
Averianov, A., MARTIN, T., Skutschas, P., Danilov, I., SCHULTZ, J.A. and SCHELLHORN, R. 2016. „Middle Jurassic vertebrate assemblage of Berezovsk coal mine in western Siberia (Russia)“. Global Geology. 2016(19(4)): 187-204.
https://doi.org/10.3969/j.issn.1673-9736.2016.04.01
Luo, Z.-X., Schultz, J.A. & Ekdale, E. G. 2016. „Chapter 6. Evolution of the middle and inner ears of mammaliaforms: The approach to mammals“. In: Evolution of the vertebrate ear: A paleontological perspective. 139-174. J. A. Clack, R. R. Fay and A. N. Popper (eds.). Springer Handbook of Auditory Research.
Martin, T., Schultz, J.A., Schwermann, A.H. and Wings, O. 2016. „First Jurassic Mammals of Germany: Multituberculate teeth from Langenberg Quarry (Lower Saxony)“. Acta Palaeontologica Polonica. 67(67): 171-179.
https://doi.org/10.4202/pp.2016.67_171
Schultz, J. A., Zeller U. & Luo Z. X. 2016. „Inner ear labyrinth anatomy of monotremes and implications for mammalian inner ear evolution“. Journal of Morphology. 278(2): 236-263.
Hielscher, R.C., Schultz, J.A. and Martin, T. 2015. „Wear pattern of the molar dentition of an extant and an Oligocene bat assemblage with implications on functionality“. Palaeobiodiversity and Palaeoenvironments. 95(4): 597-611.
https://doi.org/10.1007/s12549-015-0186-z
Schultz J. A., Kause, D.W., Koenigswald, W. v. 2015. „Dental function and diet of Vintana sertichi (Mammalia, Gondwanatheria) from the Late Cretaceous of Madagascar“. Journal of Vertebrate Paleontology. 34, Memoir 1: 182-202.
Mähler B., A. H. Schwermann, M. Wuttke, J. A. Schultz & T. Martin 2015. „4D-Virtopsy and the taphonomy of a mole from the Oligocene of Lake Enspel (Germany)“. Palaeobiodiversity and Palaeoenvironments. 95: 115-131.
Luo Z.-X., J. A. Schultz & E. G. Ekdale 2015. „Chapter 6. Evolution of the middle and inner ears of mammaliaforms: Perspectives from the fossil record“. In: Evolution of the Vertebrate Ear: A Paleontological Perspective. pp 139-174. eds. J. A. Clack, R. R. Fay and A. N. Popper. Springer. ISBN: 978-3-319-46659-0.
Krause, D.W., Hoffmann, S., Wible, J.R., Kirk, E.C., Schultz, J.A., Koenigswald, v. W., Groenke, J.R., Rossie, J.B., O’Connor, P.M., Seiffert, E.R., Dumont, E.R., Holloway, W.L., Rogers, R.R., Rahantarisoa, L.J., Kemp, A.D. and Andriamialison, H. 2014. „First cranial remains of a gondwanatherian mammal reveal remarkable mosaicism“. Nature. 515(7528): 512-517.
https://doi.org/10.1038/nature13922
Schultz, J.A. and Martin, T. 2014. „Function of pretribosphenic and tribosphenic mammalian molars inferred from 3D animation“. Naturwissenschaften. 101(10): 771-781.
https://doi.org/10.1007/s00114-014-1214-y
Angelone,C., Schultz,J. and Erbajeva,M. 2014. „Determining the ontogenetic variation of lower cheek teeth occlusal surface patterns in lagomorphs using micro-ct technology“. Palaeontologia Electronica. (): .
https://doi.org/10.26879/415
Koenigswald, W.v. , Anders, U., Engels, S., Schultz, J.A. and Kullmer, O. 2013. „Jaw movement in fossil mammals: analysis, description and visualization“. Paläontologische Zeitschrift. 87(1): 141-159.
https://doi.org/10.1007/s12542-012-0142-4
Hoffmann,R., Schultz,J.A., Schellhorn,R., Rybacki,E., Keupp,H., Gerden,S.R., Lemanis,R. and Zachow,S. 2013. „Non-invasive imaging methods applied to neo- and paleontological cephalopod research“. . (): .
https://doi.org/10.5194/bgd-10-18803-2013
Schultz, J. A. 2012. „Funktionelle Morphologie und Abnutzungsmuster prätribosphenischer Molaren am Beispiel der Dryolestida (Mammalia, Cladotheria)“. Dissertation Rheinische Friedrich-Wilhelms-Universität Bonn. 170 pp.http://hss.ulb.uni-bonn.de/2012/2787/2787.htm .
Luo, Z., Ruf, I., Schultz, J.A. and Martin, T. 2011. „Fossil evidence on evolution of inner ear cochlea in Jurassic mammals“. Proceedings of the Royal Society B: Biological Sciences. 278(1702): 28-34.
https://doi.org/10.1098/rspb.2010.1148
Schultz, J.A. and Martin, T. 2011. „Wear pattern and functional morphology of dryolestoid molars (Mammalia, Cladotheria)“. Paläontologische Zeitschrift. 85(3): 269-285.
https://doi.org/10.1007/s12542-010-0091-8
Koenigswald, W.v., Anders, U., Engels, S., Schultz, J.A. and Ruf, I. 2010. „Tooth Morphology in Fossil and Extant Lagomorpha (Mammalia) Reflects Different Mastication Patterns“. Journal of Mammalian Evolution. 17(4): 275-299.
https://doi.org/10.1007/s10914-010-9140-z
Lazzari, V., Schultz, J.A., Tafforeau, P. and Martin, T. 2010. „Occlusal Pattern in Paulchoffatiid Multituberculates and the Evolution of Cusp Morphology in Mammaliamorphs with Rodent-like Dentitions“. Journal of Mammalian Evolution. 17(3): 177-192.
https://doi.org/10.1007/s10914-010-9139-5

Awards and Grants

2023 DAAD Funding of congress participation for educators "International Congress of Vertebrate Morphology" in Cairns Australia
2019 DAAD Funding of congress participation for educators "Annual Meeting of the Society of Vertebrate Palaeontology" in Brisbane, Australia
2015 DAAD Postdoc fellowship, two year research funding in the US
2013 Poster Award of the Paläontologische Gesellschaft
2012 Tilly-Edinger-Award for young academics of the Paläontologische Gesellschaft
2009 Project EC-440 at the ESRF (European Synchrotron Radiation Facility) in Grenoble, France

Fieldwork experiences

Excavation; Middle Jurassic vertebrate fossil remains (Bathonian, Itat Fm.) from the Berezovsk Quarry of West-Sibiria, Russia (Collaboration Universität Bonn and Zoological Insitute of the Russian Academy of Science).
Excavation; Early Cretaceous vertebrate fossil remains (Cenomanian, Mata Amarilla Fm.) in Patagonia, South America (Collaboration Universität Bonn and Museo de La Plata.
Field assistant for ecological study of small mammal behavior in arid habitats in Namibia (Collaboration Humboldt Universität Berlin and Universität Köln)
Field training for collecting and quarrying fossils from the late Oligocene oil shale sediments of the Ölschiefergrube Enspel, (Germany).

Editorial & Review Activities

Since 2021 Editorial Board of the Journal of Mammalian Evolution
Reviews for different Journals (Nature, Science, Zoological Journal of the Linnean Society, Plos1, Journal of Morphology, Journal of Anatomy, Journal of Biomechanics, Journal of Vertebrate Paleontology among others)

Memberships and Board Activities

Since 2022 Vice President and Advisory Board Gesellschaft für Biologische Systematik (GfBS)
Since 2019 Advisory Board Paläontologische Gesellschaft (PalGes)
Since 2019 Member of the Gesellschaft für Biologische Systematik (GfBS)
Since 2008 Member of the Paläontologische Gesellschaft (PalGes)
Since 2008 Member of the Society of Vertebrate Paleontology (SVP)

Miscellaneous

Research student in the DFG Graduiertenkolleg 503 “Evolutionary Transformations and Mass Extinctions” of the Humboldt Universität zu Berlin (Germany); Project Title: Comparative investigations of the neonates of the golden hamster (Mesocricetus auratus), the grey short-tailed opossum (Monodelphis domestica) and the short-eared elephant shrew (Macroscelides proboscideus). Link: https://gepris.dfg.de/gepris/projekt/272675?language=en

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